Edward Stanley
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ABOUT ME

Education:
2009-2013 Richard Gilder Graduate School, American Museum of Natural History, New York, NY
Ph.D. Comparative Biology. Systematics and morphological diversification of the Cordylidae (Squamata)
2007-2009 Villanova University, Villanova, PA
M.S. Biology. Evolutionary relationships within the family Cordylidae (Squamata)
2003-2004 University of Richmond, Richmond, Virginia
Student exchange program
2001-2005 University of St Andrews, St. Andrews, Scotland
B.Sc Hons. Zoology
Professional Preparation:
Director, Digital Discovery and Dissemination (3D) Laboratory, Florida Museum of Natural History, Gainesville, FL. February 2018-present
Associate Scientist, Florida Museum of Natural History, Gainesville, FL. July 2017 – present
Postdoctoral researcher, University of Florida, Gainesville, FL.
October 2015 – July 2017
Postdoctoral researcher, California Academy of Sciences,
San Francisco, CA. October 2013 – October 2015
Research Associate, California Academy of Sciences, San Francisco, CA. September 2015–present
Research Associate, American Museum of Natural History, New York, NY. August October 2013 – present
Associate Editor, African Journal of Herpetology.
December 2014–present

Curriculum Vitae
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Publications :

2022
Čerňanský, A., Stanley, E.L., Daza, J.D., Bolet, A., Arias, J.S., Bauer, A.M., Vidal-García, M., Bevitt, J.J., Peretti, A.M., Aung, N.N. and Evans, S.E. 2022. A New Early Cretaceous lizard in Myanmar Amber with Exceptionally Preserved Integument. Scientific Reports 12, 1660 
Open Access: Download here

2021
Paluh, D.J., Dillard, W.A., Stanley, E.L., Fraser, G.J. and Blackburn, D.C., 2021. Re‐evaluating the morphological evidence for the re‐evolution of lost mandibular teeth in frogs. Evolution.
Open Access: Download here
Pinheiro, P.D., Blotto, B.L., Ron, S.R., Stanley, E.L., Garcia, P.C., Haddad, C.F., Grant, T. and Faivovich, J., 2021. Prepollex diversity and evolution in Cophomantini (Anura: Hylidae: Hylinae). Zoological Journal of the Linnean Society.
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Bell, C. J., Daza, J. D., Stanley, E. L., & Laver, R. J. 2021. Unveiling the elusive: X-rays bring scolecophidian snakes out of the dark. The Anatomical Record, 1– 8. https://doi.org/10.1002/ar.24729
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Laver, R.J., Daza, J.D., Ellis, R.J., Stanley, E.L. and Bauer, A.M., 2021. Underground Down Under: Skull anatomy of the southern blind snake Anilios australis Gray, 1845 (Typhlopidae: Serpentes: Squamata). The Anatomical Record.
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Bolet, A.; Stanley, E. L.; Daza, J. D.; Salvador Arias, J.; Čerňanský, A.; Vidal-García, M.; Bevitt, J. J.; Peretti, A. and Evans, S. E. (2021). "Unusual morphology in the mid-Cretaceous lizard Oculudentavis". Current Biology, 31, 1–12. doi:10.1016/j.cub.2021.05.040.
Open Access: Download here
Paluh, D.J., Riddell, K., Early, C.M., Hantak, M.M., Jongsma, G.F., Keeffe, R.M., Silva, F.M., Nielsen, S.V., Vallejo-Pareja, M.C., Stanley, E.L. and Blackburn, D.C., 2021. Rampant tooth loss across 200 million years of frog evolution. Elife, 10, p.e66926.
Open Access: Download here
Blackburn, D.C., Hernández-Morales, C., Paluh, D.J. and Stanley, E.L., 2021. Predation on a gymnophthalmid lizard by the Brazilian Dumpy Frog, Stereocyclops incrassatus (Microhylidae, Gastrophryninae). Herpetology Notes, 14, pp.847-849.
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Bardua, C., Fabre, A., Clavel, J.,  Bon, M.B.,  Das, K., Stanley, E.L.,  Blackburn, D.C., and Goswami, A.  (2021) Size, microhabitat, and loss of larval feeding drive cranial diversification in frogs. Nat Commun 12, 2503. https://doi.org/10.1038/s41467-021-22792-y
Open Access: Download here
Goodman, C.M., Jongsma, G.F.M., Hill, J.E., Stanley, E.L., Tuckett, Q.M., Blackburn, D.C. and Romagosa, C.M. (2021) A case of mistaken identity: genetic and anatomical evidence reveal the cryptic invasion of Xenopus tropicalis in central Florida. Journal of Herpetology 55.1:62-69.
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Wagner, P., Stanley, E.L., Daza, J.D., Bauer, A.M. (2021) A new agamid lizard in mid Cretaceous amber from northern Myanmar. Cretaceous Research
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Feiner, N., Jackson, I.S.C., Stanley, E.L., and Tobias Uller, T. (2021) Evolution of the locomotor skeleton in Anolis lizards reflects the interplay between ecological opportunity and phylogenetic inertia. Nature Communications 12, 1525
Open Access: Download here

2020
Daza J.D., Stanley, E.L., Bolet, A,, Bauer, A.M., Salvadore Arisa, J.m Cernasky, A., Bevitt, J., Wagner, P., and Evans, S.E. (2020) Enigmatic amphibians in mid-Cretaceous amber were chameleon-like and had ballistic feeding. Science. 370(6517): 687-691.
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Nielsen SV, Conradie W, Ceríaco LMP, Bauer AM, Heinicke MP, Stanley EL, Blackburn DC (2020) A new species of Rain Frog (Brevicipitidae, Breviceps) endemic to Angola. ZooKeys 979: 133-160. https://doi.org/10.3897/zookeys.979.56863
Open Access: Download here

Blackburn, D.C
., Nielsen, S.V., Barej, M.F., Hirschfeld, J.D.M., Kouamé N.G., Lawson D.,  Loader, S.,  Ofori‐Boateng, C., Stanley, E.L. and Rödel MO. 2020. Evolution of the African slippery frogs (Anura: Conraua), including the world’s largest living frog. Zoolica Scripta. 2020; 00: 1– 13.
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Heinicke M.P., Titus-McQuillan, J.E., Daza, J.D., Kull, E.M., Stanley, E.L. and  Bauer, A.M. 2020.Phylogeny and evolution of unique skull morphologies in dietary specialist African shovel-snouted snakes (Lamprophiidae: Prosymna), Biological Journal of the Linnean Society, , blaa076.
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Fabre, A.-C., C. Bardua, M. Bon, J. Clavel, R.N. Felice, J.W. Streicher, J. Bonnel, Stanley, E.L., D.C. Blackburn and A. Goswami. 2020. Metamorphosis and the evolution of morphological diversity in salamanders. Nature Ecology & Evolution 2020.
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Bardua, C., Fabre, A.‐C., Bon, M., Das, K., Stanley, E.L., Blackburn, D. and Goswami, A. 2020, Evolutionary integration of the frog cranium. Evolution. doi:10.1111/evo.13984
Open Access: Download here
Paluh, D.P., Stanley, E.L. and Blackburn, D.C., 2020 Evolution of hyperossification expands skull diversity in frogs. Proceedings of the National Academy of Sciences. 117 (15) 8554-8562
Open Access: Download here
Bates, M.F. and Stanley, E.L. 2020 A taxonomic revision of the south-eastern dragon lizards of the Smaug warreni (Boulenger) species complex in southern Africa, with the description of a new species (Squamata: Cordylidae). PeerJ 8:e8526
Open Access: Download here
Singer, R., Moore, J. and Stanley, E.L. 2020 Novel life history strategy in a deep sea fish challenges assumptions about reproduction in extreme environments. Scientific Reports 10, 3649
Open Access: Download here
Claunch N., Stanley E.L. and Romagosa C.M. 2020 Natural History note: Leiocepahalus carinatus (Northern Curly-Tailed Lizard) Extreme Fecal Impaction. Herpetological Review 51(1): p126.
Open Access: Download here

2019
Blackburn, D.C., L. Roberts, M.C. Vallejo-Pareja, and. Stanley, E.L.. 2019. First record of the anuran family Rhinophrynidae from the Oligocene of eastern North America. Journal of Herpetology​. Vol. 53, No. 4, pp. 316-323.
Open Access: Download here
Blackburn D.C., Giribet G., Soltis D.G. and Stanley, E.L. 2019. Predicting the Impact of Describing New Species on Phylogenetic Patterns. Integrative Organismal Biology, obz 028
Open Access: Download here
Černanský A., Stanley E.L. 2019. The atlas
– axis complex in Dibamidae (Reptilia: Squamata) and their potential relatives: The effect of a fossorial lifestyle on the morphology of this skeletal bridge. Journal of Morphology. 2019;1–21.
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Marques, M., Ceríaco, L.M.P., Stanley, E.L., Bandeira, S.A., Agarwal, I. and Bauer A.M., 2019. A new species of Girdled Lizard (Squamata: Cordylidae) from the Serra da Neve Inselberg, Namibe Province, southwestern Angola. Zootaxa 4668(4)
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Evans B.J., Gansauge, M-T., Stanley, E.L., Furman, B.L.S., Caroline M. S. Cauret, C.M.S., Ofori-Boateng, C., Gvoždík, V., Streicher, J.W., Greenbaum, E., Tinsley, R.C., Meyer, M. and Blackburn D.C. 2019. Xenopus fraseri: Mr. Fraser, where did your frog come from? PLoS ONE 14(9): e0220892. https://doi.org/10.1371/journal.pone.0220892
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Holovacs, N.T., Daza, J.D., Guerra, C., Stanley, E.L. and Montero, R., 2019. You can't run, but you can hide: the skeleton of the sand‐swimmer lizard Calyptommatus leiolepis (Squamata: Gymnophthalmidae). The Anatomical Record.
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Griffing A.H., Sanger T.J., Daza J.D., Nielsen S.V., Pinto B.J., Stanley E.L., and Gamble T. 2019 Embryonic development of a parthenogenetic vertebrate, the mourning gecko . Embryonic development of a parthenogenetic vertebrate, the mourning gecko (Lepidodactylus lugubris). Developmental Dynamics. 2019; 248: 1070– 1090.
doi:10.1002/dvdy.72
Camaiti, M. , Villa, A. , Wencker, L. C., Bauer, A. M., Stanley, E. L. and Delfino, M. 2019, Descriptive osteology and patterns of limb loss of the European limbless skink Ophiomorus punctatissimus (Squamata, Scincidae). Journal of Anatomy. 235: 313-345.
doi:10.1111/joa.13017
Blackburn D.C., Paluh, D.J., Krone, I., Roberts, E.M., Stanley, E.L. and Stevens, N.J. "The Earliest Fossil of the African Clawed Frog (Genus Xenopus) from Sub-Saharan Africa," Journal of Herpetology, 53(2), 125-130.
Open Access: Download here
Portillo F.,  Stanley E.L.,   Branch W.R.,   Conradie W.R., Rödel M.O.,   Penner J.,  Barej M.,  Kusamba, C., Muninga W.M., Aristote M.M.,  Bauer A.M., Trape J.F.,  Nagy Z.T.,  Carlino P.,  Pauwels O.S.G.,  Menegon M.,  Ineich I.,  Burger M.,  Mazuch T., Jackson K.;  Hughes D.F., Behangana M and Greenbaum E. 2019. Evolutionary history of burrowing asps (Atractaspidinae) (Squamata: Lamprophiidae) with emphasis on fang evolution and prey selection PLoS One 14 (4) doi.org/10.1371/journal.pone.0214889
Open Access: Download here
Vijayakumar S.P., Pyron, R.A., Dinesh K.P., Torsekar V.R., Srikanthan A.N.,  Swamy P.,  Stanley E.L.,  Blackburn D.C.,  Shanker K. 2019. A new ancient lineage of frog (Anura: Nyctibatrachidae: Astrobatrachinae subfam. nov.) endemic to the Western Ghats of Peninsular India, PeerJ 7:e6457 https://doi.org/10.7717/peerj.6457
Open Acess: Download here

2018
Daza
, J. D.,   Bauer, A. M.,   Stanley, E. L.,  Bolet, A., Dickson, B., and  Losos, J. B. 2018. An Enigmatic Miniaturized and Attenuate Whole Lizard from the Mid-Cretaceous Amber of Myanmar Breviora 563 (1), 1-18
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Ceríaco, L.M.P.,  Marques, M.P., Bandeira, S.,  Agarwal, I., Stanley, E.L., Bauer, A.M., Heinicke, M.P.,  Blackburn, D.C. 2018. A new earless species of Poyntonophrynus (Anura, Bufonidae) from the Serra da Neve Inselberg, Namibe Province, Angola. ZooKeys 780: 109–136, doi: 10.3897/zookeys.780.25859
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Xing, L., Stanley, E.L., Bai, M.
and Blackburn, D.C. 2018. The earliest direct evidence of frogs in wet tropical forests from Cretaceous Burmese amber. Scientific Reports 8, Article number: 8770
Open Access: Download here
Li Y., Ruan Y., Kasson M.T., Stanley E.L., Gillett C.P.D.T., Johnson A.J.,  Zhang M, Hulcr J. Structure of the Ambrosia Beetle (Coleoptera: Curculionidae) Mycangia Revealed Through Micro-Computed Tomography, 2018. Journal of Insect Science, Volume 18, Issue 5, September 2018, 13, https://doi.org/10.1093/jisesa/iey096
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Hill R.L., Martin, K.G., Stanley, E.L.  and  Mendelson, J.R. 2018. A taxonomic review of the genus Hemiphractus (Anura: Hemiphractidae) in Panama: Description of Two New Species, Resurrection of Hemiphractus panamensis (Stejneger, 1917), and Discussion of Hemiphractus fasciatus Peters. 1862 Zootaxa 4429 (3): 495–512
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Li, Y. , Ruan, Y. , Stanley, E. L., Skelton, J. and Hulcr, J. 2018. Plasticity of mycangia in Xylosandrus ambrosia beetles. Insect Science. 00, 0-11. doi:10.1111/1744-7917.12590
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2017
Kim, B.G., Kim, Y.H., Stanley, E.L., Garrido-Martin, E.M.,  Lee, Y.J.,  Oh, S.P. 2017. CXCL12-CXCR4 signaling plays an essential role in proper patterning of aortic arch and pulmonary arteries, Cardiovascular Research, , cvx188,
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Ribeiro, L. F., Blackburn, D. C., Stanley, E. L., Pie, M. R., and Bornschein, M. R. 2017. Two new species of the Brachycephalus pernix group (Anura: Brachycephalidae) from the state of Paraná, southern Brazil. PeerJ, 5, e3603.
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Blackburn D.C.,  Paluh, D.J. and Stanley, E.L. 2017. Phrynobatrachus krefftii (Krefft’s Puddle Frog). Anurophagy. Herpetological Review 48: 413–414.
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Bateman,C.,  Yin-Tse Huang, Simmons, D.R., Kasson, M.T., Stanley, E.L., and Hulcr, J. 2017 Ambrosia beetle Premnobius cavipennis (Scolytinae: Ipini) carries highly divergent ascomycotan ambrosia fungus, Afroraffaelea ambrosiae gen. nov. sp. nov. (Ophiostomatales). Fungal Ecology, 25: 41–49
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2016
Bornschein, M.R., Ribeiro, L.F., Blackburn, D.C., Stanley, E.L. and Pie, M.R. 2016. A new species of Brachycephalus (Anura: Brachycephalidae) from Santa Catarina, southern Brazil. PeerJ, 4 e2629.
Open Access: Download here
Blackburn, D.C., Boix, C., Greenbaum, E., Fabrezi, M., Meirte, D., Plumptre, A.J. and Stanley, E.L. 2016. The distribution of the Bururi Long-fingered Frog (Cardioglossa cyaneospila, family Arthroleptidae), a poorly known Albertine Rift endemic. Zootaxa 4170 (2)
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Ceríaco, L.M.P., de Sá, S.C., Bandeira, S., Valério, H., Stanley, E.L., Kuhn, A.L., Marques, M., Vindum, J.V.,  Blackburn D.C., and Bauer, A.M. 2016. Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a Synoptic list of the Amphibians and Reptiles of Namibe Province, Southwestern Angola. Proceedings of the California Academy of Sciences 63 (2)
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Daza, J.D., Stanley, E.L., Wagner, P., Bauer, A.M. and Grimaldi, D.A. 2016. Mid-Cretaceous amber fossils illuminate the past diversity of tropical lizards. Science Advances 2(3)
​Open Access: Download here

Stanley, E.L., Ceríaco, L.M.P., Bandeira, S. Valerio, H., Bates, M.F., and Branch,W.R. 2016. A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert. Zootaxa 4061(3): 201–226
Open Access: Download here

2015
Evans BJ, Carter TF, Greenbaum E, Gvoždík V, Kelley, D.B., McLaughlin, P.J., Pauwels, O.S.G., Portik, D.M., Stanley, E.L., Tinsley, R.C.,Tobias, M.L., Blackburn, D.C.
2015. Genetics, Morphology, Advertisement Calls, and Historical Records Distinguish Six New Polyploid Species of African Clawed Frog (Xenopus, Pipidae) from West and Central Africa. PLoS ONE 10(12): e0142823. doi: 10.1371/journal.pone.0142823
Open Access: Download here

2014
Stanley, E.L
, and Bates, M.F. 2014. Here be Dragons: A phylogenetic and biogeographic study of the Smaug warreni species complex (Squamata: Cordylidae) in southern Africa. Zoological Journal of the Linnean Society 172: 892­­­–909
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de Sá R.O., Grant, T.A., Camargo,  A., Heyer, W.R., Ponssa, M.L. and Stanley E.L. 2014. Systematics of the Neotropical Genus LeptodactylusFitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the Relevance of Non-molecular Evidence, and Species Accounts. South American Journal of Herpetology 9:S1–100
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Stanley, E.L. Cordylid systematics and evolution. 2014. In Girdled Lizards and their relatives. Natural history, captive care and breeding.  Reissig, J. Edition Chimaira London p.249.
Find it here

2012
Mouton, P.F.N., Fleming, A. and Stanley, E. L., 2012. Synchronised versus asynchronised breeding in the lizard family Cordylidae: An evolutionary perspective. Journal of Zoology  288:191–198
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Greenbaum, E.B., Stanley, E.L., Kusamba, C., Moninga W.M., Goldberg, S.R., and Bursey, C.R. 2012. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of southeastern Democratic Republic of the Congo. African Journal of Herpetology. 61:14-39
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Portik D, Wood P, Grismer J, Stanley E.L. and Jackman T. 2012. Identification of 104 rapidly-evolving nuclear protein-coding markers for amplification across scaled reptiles using genomic resources. Conservation Genetics Resources 4: 1-10
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2011
Stanley, E. L., Bauer, A. M., Jackman, T. R., Mouton, P. L. N. and Branch, W. R. 2011. Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizards (Cordylidae: Squamata). Molecular Phlylogenetics and Evolution 58: 53-70
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2008
Stanley, E. L.
, Kendal, R. L., Kendal, J. R., Grounds S., and Laland K. N. 2008. The effects of group size, rate of turnover and disruption to demonstration on the stability of foraging traditions in fish. Animal Behaviour 75: 565-572
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Patents:

Scheiffele, G. and Stanley, E.L., (2021). Methods and compositions for minimizing X-ray scattering artifact (US Patent No.11,183,314)

Grants and Awards:

National Science Foundation DMR#2017977, $1,202,863. MRI: Acquisition of Nano-resolution Zeiss Xradia 620 Versa X-ray Computed Tomography with Phase Contrast and Lab Diffraction Contrast Tomography, Co-Principal Investigator, July 2020

National Science Foundation ICB#1929448, $763,978. CSBR: Curating the Early Anthropocene Record of circum-Caribbean Animal Biodiversity in the Florida Museum of Natural History, Co-Principal Investigator, September 2019

National Science Foundation SBS #1924759 $104,252. Collaborative Research: Repeated Adaptive Radiation in the Coevolutionary History of Birds and Feather Lice, Principal Investigator, September 2019.

National Science Foundation DBI #1701714 $808,235. Digitization TCN: Collaborative Research: oVert: Open Exploration of Vertebrate Diversity in 3D, Co-Principal Investigator, September 2017. NSF page. iDigBio Wiki page.

National Science Foundation DEB #1556559
. $884,855. Collaborative Research: Survey and Analysis of the Angolan Herpetofauna. Senior personnel, June 2016. NSF page.

National Science Foundation DEB #1210478. $13,910. Doctoral Dissertation Improvement Grant:  A synthetic analysis of evolutionary patterns in the Squamate family Cordylidae. Co-Principal Investigator, March 2012. NSF page.

Exploration Fund grant, the Explorers Club. $1050 for the project “Phylogenetic relationships of the Cape cliff lizards.” May 2011

Tracy Storer award, Joint meeting of Ichthyologists and Herpetologists, Providence, RI, July 2010.

Full-year Research Fellowship, Villanova University, Villanova, PA, Sept 2008 to May 2009.

Summer Research Fellowship, Villanova University, Villanova, PA, June to July 2008.

Nesmith Dickinson Research Award, University of Richmond, Richmond, VA 2004. Awarded to a student majoring in biology at the University of Richmond to aid research projects during the recipient's senior year.

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  • Home
  • Research Interests
    • CT scanning >
      • CT processing
    • Girdled lizards
  • About Me
  • Contact and links